The evolutionary ancestry of arthropods dates back to the Cambrian period. The group is generally regarded as monophyletic, and many analyses support the placement of arthropods with cycloneuralians (or their constituent clades) in a superphylum Ecdysozoa. Overall, however, the basal relationships of animals are not yet well resolved. Likewise, the relationships between various arthropod groups are still actively debated. Today, arthropods contribute to the human food supply both directly as food, and more importantly, indirectly as pollinators of crops. Some species are known to spread severe disease to humans, livestock, and crops. (Full article...)
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Hemigrapsus estellinensis is an extinctspecies of crab, formerly endemic to the Texas Panhandle. It was discovered by Gordon C. Creel in 1962 and was probably already extinct before his description was published in 1964, after the Estelline Salt Springs where it lived were contained by the United States Army Corps of Engineers. H. estellinensis is closely related to species from the Pacific Ocean such as Hemigrapsus oregonensis, but lived 800 km (500 mi) inland in a hypersaline spring. It differed from its relatives by the pattern of spots on its back, and by the relative sizes of its limbs. (Full article...)
Chelicerata split from Mandibulata by the mid-Cambrian, as evidenced by stem-group chelicerates like Habeliida and Mollisonia present by this time. The surviving marine species include the four species of xiphosurans (horseshoe crabs), and possibly the 1,300 species of pycnogonids (sea spiders), if the latter are indeed chelicerates. On the other hand, there are over 77,000 well-identified species of air-breathing chelicerates, and there may be about 500,000 unidentified species.
Like all arthropods, chelicerates have segmented bodies with jointed limbs, all covered in a cuticle made of chitin and proteins. The chelicerate body plan consists of two tagmata, the prosoma and the opisthosoma, except that mites have lost a visible division between these sections. The chelicerae, which give the group its name, are the only appendages that appear before the mouth. In most sub-groups, they are modest pincers used to feed. However, spiders' chelicerae form fangs that most species use to inject venom into prey. The group has the open circulatory system typical of arthropods, in which a tube-like heart pumps blood through the hemocoel, which is the major body cavity. Marine chelicerates have gills, while the air-breathing forms generally have both book lungs and tracheae. In general, the ganglia of living chelicerates' central nervous systems fuse into large masses in the cephalothorax, but there are wide variations and this fusion is very limited in the Mesothelae, which are regarded as the oldest and most basal group of spiders. Most chelicerates rely on modified bristles for touch and for information about vibrations, air currents, and chemical changes in their environment. The most active hunting spiders also have very acute eyesight. (Full article...)
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Calliphora vomitoria, known as the blue bottle fly, orange-bearded blue bottle, or bottlebee, is a species of blow fly, a species in the family Calliphoridae. Calliphora vomitoria is the type species of the genusCalliphora. It is common throughout many continents including Europe, Americas, and Africa. They are fairly large flies, nearly twice the size of the housefly, with a metallic blue abdomen and long orange setae on the gena.
While adult flies feed on nectar, females deposit their eggs on rotting corpses, making them important forensic insects, as their eggs and timing of oviposition can be used to estimate time of death. (Full article...)
Entomologists reserve the term bug for Hemiptera or Heteroptera, which does not include other arthropods or insects of other orders such as ants, bees, beetles, or butterflies. In some variations of English, all terrestrialarthropods (including non-insect arachnids, and myriapods) also fall under the colloquial understanding of bug.
Unionopterus was described in 1948 after the discovery of its only known specimen in the Karaganda Formation of Kazakhstan (at that time part of the Soviet Union). This was one of the first discoveries of eurypterids in the Soviet Union and Carboniferous eurypterids in general. It was probably a swimming organism as well as other eurypterids, although not as good as its relatives.
There are numerous factors that have made Unionopterus a problematic genus for eurypterid researchers. It has been placed in the familyAdelophthalmidae and it has even been speculated that a species of Adelophthalmus, A. dumonti, actually belongs to Unionopterus, but this cannot be confirmed. Many authors have chosen to completely ignore the genus during phylogenetic studies, making Unionopterus an enigmatic eurypterid. (Full article...)
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Throughout history, spiders have been depicted in popular culture, mythology and in symbolism. From Greek mythology to African folklore, the spider has been used to represent a variety of things, and endures into the present day with characters such as Shelob from The Lord of the Rings and Spider-Man from the eponymous comic series. It is also a symbol of mischief and malice for its toxic venom and the slow death it causes, which is often seen as a curse. In addition, the spider has inspired creations from an ancient geoglyph to a modern steampunkspectacle. Spiders have been the focus of fears, stories and mythologies of various cultures for centuries.
The spider has symbolized patience and persistence due to its hunting technique of setting webs and waiting for its prey to become ensnared. Numerous cultures attribute the spider's ability to spin webs with the origin of spinning, textile weaving, basket weaving, knotwork and net making. Spiders are associated with creation myths because they seem to weave their own artistic worlds. Philosophers often use the spider's web as a metaphor or analogy, and today terms such as the Internet or World Wide Web evoke the inter-connectivity of a spider web.
The external morphology of Lepidoptera is the physiological structure of the bodies of insects belonging to the order Lepidoptera, also known as butterflies and moths. Lepidoptera are distinguished from other orders by the presence of scales on the external parts of the body and appendages, especially the wings. Butterflies and moths vary in size from microlepidoptera only a few millimetres long, to a wingspan of many inches such as the Atlas moth. Comprising over 160,000 described species, the Lepidoptera possess variations of the basic body structure which has evolved to gain advantages in adaptation and distribution.
Lepidopterans undergo complete metamorphosis, going through a four-stage life cycle: egg, larva or caterpillar, pupa or chrysalis, and imago (plural: imagines) / adult. The larvae – caterpillars – have a toughened (sclerotised) head capsule, chewing mouthparts, and a soft body, that may have hair-like or other projections, three pairs of true legs, and up to five pairs of prolegs. Most caterpillars are herbivores, but a few are carnivores (some eat ants, aphids, or other caterpillars) or detritivores. Larvae are the feeding and growing stages and periodically undergo hormone-induced ecdysis, developing further with each instar, until they undergo the final larval–pupal moult. The larvae of many lepidopteran species will either make a spun casing of silk called a cocoon and pupate inside it, or will pupate in a cell under the ground. In many butterflies, the pupa is suspended from a cremaster and is called a chrysalis.
The adult body has a hardened exoskeleton, except for the abdomen which is less sclerotised. The head is shaped like a capsule with appendages arising from it. Adult mouthparts include a prominent proboscis formed from maxillary galeae, and are adapted for sucking nectar. Some species do not feed as adults, and may have reduced mouthparts, while others have them modified for piercing and suck blood or fruit juices. Mandibles are absent in all except the Micropterigidae which have chewing mouthparts. Adult Lepidoptera have two immobile, multi-faceted compound eyes, and only two simple eyes or ocelli, which may be reduced. The three segments of the thorax are fused together. Antennae are prominent and besides the faculty of smell, also aid navigation, orientation, and balance during flight. In moths, males frequently have more feathery antennae than females, for detecting the female pheromones at a distance. There are two pairs of membranouswings which arise from the mesothoracic (middle) and metathoracic (third) segments; they are usually completely covered by minute scales. The two wings on each side act as one by virtue of wing-locking mechanisms. In some groups, the females are flightless and have reduced wings. The abdomen has ten segments connected with movable inter-segmental membranes. The last segments of the abdomen form the external genitalia. The genitalia are complex and provide the basis for family identification and species discrimination. (Full article...)
Bees are found on every continent except Antarctica, in every habitat on the planet that contains insect-pollinated flowering plants. The most common bees in the Northern Hemisphere are the Halictidae, or sweat bees, but they are small and often mistaken for wasps or flies. Bees range in size from tiny stingless bee species, whose workers are less than 2 millimetres (0.08 in) long, to Megachile pluto, the largest species of leafcutter bee, whose females can attain a length of 39 millimetres (1.54 in).
Bees feed on nectar and pollen, the former primarily as an energy source and the latter primarily for protein and other nutrients. Most pollen is used as food for their larvae. Vertebrate predators of bees include primates and birds such as bee-eaters; insect predators include beewolves and dragonflies. (Full article...)
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Female with prey silver-spotted skipper (Hesperia comma)
Misumena vatia is a species of crab spider with a holarctic distribution. In North America, it is called the goldenrod crab spider or flower (crab) spider, as it is commonly found hunting in goldenrod sprays and milkweed plants. They are called crab spiders because of their unique ability to walk sideways as well as forwards and backwards. Both males and females of this species progress through several molts before reaching their adult sizes, though females must molt more to reach their larger size. Females can grow up to 10 mm (0.39 in) while males are quite small, reaching 5 mm (0.20 in) at most. Misumena vatia are usually yellow or white or a pattern of these two colors. They may also present with pale green or pink instead of yellow, again, in a pattern with white. They have the ability to change between these colors based on their surroundings through the molting process. They have a complex visual system, with eight eyes, that they rely on for prey capture and for their color-changing abilities. Sometimes, if Misumena vatia consumes colored prey, the spider itself will take on that color.
Misumena vatia feed on common insects, often consuming prey much larger than themselves. They use venom to immobilize their prey, though they are harmless to humans. They face threats due to parasites and larger insects. For Misumena vatia, survival depends on the choice of hunting site. The spiders closely monitor multiple sites to see if others nearby are frequented by greater numbers of potential prey. The primary sex ratio is biased toward females. Females are stationary and choose a flower to settle on, while males cover great distances searching for mates. Females do not emit pheromones, rather, they leave "draglines" of silk behind them as they move, which males follow. Females live longer than males, on average. After mating, females guard their nests until the young have hatched, after which they die. (Full article...)
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Interpretive drawing of the holotype and only known specimen of Dorfopterus, including a close-up on the ornamentation of the fossil.
The only known specimen of Dorfopterus consists of an incomplete uncrushed telson (the posteriormost division of the body), which was long, narrow, styliform and with a central carina ("keel"). It had a special ornamentation consisting of rib-like curved lines on each side of the telson with reticulated (net-like) patterns between them. This ornamentation was unique, and was not known to have occurred in any other eurypterid at the time of the specimen's discovery.
Originally described as part of Stylonuridae by the American paleontologist Erik Norman Kjellesvig-Waering in 1955, the strange morphology and the little known fossil material of Dorfopterus have made the classification of this genus problematic. Currently, it is considered as incertae sedis (a taxon with unclear relationships) within Eurypterida, although Dorfopterus being a eurypterid at all has also been questioned. The locality in which Dorfopterus was found, the Beartooth Butte Formation, is home to fossils of many fish, plants and a few other eurypterids. The genus is believed to have lived in an estuarine inland channel. (Full article...)
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Yellow morph of J. lagostoma
Johngarthia lagostoma is a species of terrestrial crab that lives on Ascension Island and three other islands in the South Atlantic. It grows to a carapace width of 110 mm (4.3 in) on Ascension Island, where it is the largest native land animal. It exists in two distinct colour morphs, one yellow and one purple, with few intermediates. The yellow morph dominates on Ascension Island, while the purple morph is more frequent on Rocas Atoll. The species differs from other Johngarthia species by the form of the third maxilliped.
Johngarthia lagostoma lives in burrows among vegetation, at altitudes of up to 400 m (1,300 ft), emerging at night to feed on plant matter and occasionally on animals. From January to March there is an annual migration to the sea to release the planktoniclarvae. The species was first described (as Gecarcinus lagostoma) by Henri Milne-Edwards in 1837 from material sent to him by the naturalists Jean René Constant Quoy and Joseph Paul Gaimard, collected by the French ship Astrolabe. (Full article...)
Butterflies are winged insects from the lepidopteransuborderRhopalocera, characterized by large, often brightly coloured wings that often fold together when at rest, and a conspicuous, fluttering flight. The group comprises the superfamiliesHedyloidea (moth-butterflies in the Americas) and Papilionoidea (all others). The oldest butterfly fossils have been dated to the Paleocene, about 56 million years ago, though they likely originated in the Late Cretaceous, about 101 million years ago.
Butterflies have a four-stage life cycle, and like other holometabolous insects they undergo complete metamorphosis. Winged adults lay eggs on the food plant on which their larvae, known as caterpillars, will feed. The caterpillars grow, sometimes very rapidly, and when fully developed, pupate in a chrysalis. When metamorphosis is complete, the pupal skin splits, the adult insect climbs out, expands its wings to dry, and flies off.
Some butterflies, especially in the tropics, have several generations in a year, while others have a single generation, and a few in cold locations may take several years to pass through their entire life cycle. (Full article...)
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Scyllarides latus, the Mediterranean slipper lobster, is a species of slipper lobster found in the Mediterranean Sea and in the eastern Atlantic Ocean. It is edible and highly regarded as food, but is now rare over much of its range due to overfishing. Adults may grow to 1 foot (30 cm) long, are camouflaged, and have no claws. They are nocturnal, emerging from caves and other shelters during the night to feed on molluscs. As well as being eaten by humans, S. latus is also preyed upon by a variety of bony fish. Its closest relative is S. herklotsii, which occurs off the Atlantic coast of West Africa; other species of Scyllarides occur in the western Atlantic Ocean and the Indo-Pacific. The larvae and young animals are largely unknown. (Full article...)
Iridomyrmex is a genus of ants called rainbow ants (referring to their blue-green iridescent sheen) first described by Austrian entomologist Gustav Mayr in 1862. He placed the genus in the subfamilyDolichoderinae of the family Formicidae. It has 79 described species and five fossil species. Most of these ants are native to Australia; others are found in Asia and Oceania, and they have been introduced to Brazil, New Zealand, and the United Arab Emirates. Fossil species are known from China, France, and the United States.
These ants are known to be an ecologically dominant and important group of ants, but they are sometimes regarded as pests because they disturb soil and enter human houses. Farmers in rural Australia place animal carcasses on meat ant (I. purpureus) mounds as a method of disposing of them; meat ants consume the carcass and reduce it to bones in a matter of weeks. Meat ants also engage in ritualised fighting, which helps prevent casualties and solve territorial disputes between neighbouring colonies. The largest members of this genus are those of the I. purpureus species group, measuring 8 mm (0.31 in).
After their nuptial flight, queen ants may establish a colony by themselves, by budding, or cooperatively, where a subset of the colony migrates to a new location or when multiple queens help find a suitable nesting spot, but they display intolerance to each other when workers are present. The eggs take 44 to 61 days to fully develop into adults. Ants of the genus live in a wide variety of habitats and nest in soil in numbers that range from a few hundred individuals to over 300,000 in a single colony. Depending on the species, nests are large mounds covered in pebbles with multiple entrances, while others live above ground in twig nests. In some cases, ants dwell in several nest sites connected by paths; some of these nests can extend to 650 m (2,130 ft) in length. Some species associate with caterpillars and butterflies that provide the ants with secretions and honeydew, and I. bicknelli pollinates orchids. These ants are predators and scavengers; they hunt for prey to feed their young. Notably, these ants are immune to the toxins of the cane toad and feed on the juveniles. Predators such as spiders, birds, lizards, and other ants prey on Iridomyrmex ants. (Full article...)
... that adult males of the parasiticwaspEncarsia perplexa can only develop when a virgin female lays eggs in a fully developed larva of her own species?
Image 2 Honeybee larvae have flexible but delicate unsclerotised cuticles. (from Arthropod exoskeleton)
Image 3 This fully-grown robber crab has tough fabric forming its joints, delicate biomineralized cuticle over its sensory antennae, optic-quality over its eyes, and strong, calcite-reinforced chitin armouring its body and legs; its pincers can break into coconuts (from Arthropod exoskeleton)
Image 4This Zoea-stage larva is hardly recognisable as a crab, but each time it sheds its cuticle it remodels itself, eventually taking on its final crab form (from Arthropod exoskeleton)
Image 10Some of the various hypotheses of myriapod phylogeny. Morphological studies (trees a and b) support a sister grouping of Diplopoda and Pauropoda, while studies of DNA or amino acid similarities suggest a variety of different relationships, including the relationship of Pauropoda and Symphyla in tree c. (from Myriapoda)
Image 18In honeypot antrepletes, the abdomens of the workers that hold the sugar solution grow vastly, but only the unsclerotised cuticle can stretch, leaving the unstretched sclerites as dark islands on the clear abdomen (from Arthropod exoskeleton)
Image 19Reconstruction of Mollisonia plenovenatrix, the oldest known arthropod with confirmed chelicerae (from Chelicerata)
Image 21Formation of anterior segments across arthropod taxa based on gene expression and neuroanatomical observations, Note the chelicera(Ch) and chelifore(Chf) arose from somite 1 and thus correspond to the first antenna(An/An1) of other arthropods. (from Chelicerata)
Image 22Ghost crab, showing a variety of integument types in its exoskeleton, with transparent biomineralization over the eyes, strong biomineralization over the pincers, and tough chitin fabric in the joints and the bristles on the legs (from Arthropod exoskeleton)
Image 28The house centipedeScutigera coleoptrata has rigid sclerites on each body segment. Supple chitin holds the sclerites together and connects the segments flexibly. Similar chitin connects the joints in the legs. Sclerotised tubular leg segments house the leg muscles, their nerves and attachments, leaving room for the passage of blood to and from the hemocoel (from Arthropod exoskeleton)
Image 29Decapods, from Ernst Haeckel's 1904 work Kunstformen der Natur (from Crustacean)
Image 33Mature queen of a termite colony, showing how the unsclerotised cuticle stretches between the dark sclerites that failed to stretch as the abdomen grew to accommodate her ovaries (from Arthropod exoskeleton)
The Ozyptila praticola species of crab spider is found throughout Europe and the Middle East. They do not build webs to trap prey, but are active hunters. Crab spiders are so named because of their first two pairs of legs, which are held out to the side giving them a crab-like appearance. Also, like crabs, these spiders move sideways and backwards more easily than forwards.
A honey bee extracts nectar from a flower using its proboscis. Tiny hairs covering the bee's body maintain a slight electrostatic charge, causing pollen from the flower's anthers to stick to the bee's hairs, allowing for pollination when the bee moves on to another flower.
The red rock crab (Grapsus grapsus), also known as "Sally Lightfoot", is one of the commonest crabs along the western seaboard of the Americas. John Steinbeck wrote of them, "Everyone who has seen them has been delighted with them ... These little crabs, with brilliant cloisonné carapaces, walk on their tiptoes, they have remarkable eyes and an extremely fast reaction time." He tried to catch them but to little avail. "If you walk slowly, they move slowly ahead of you in droves. If you hurry, they hurry. When you plunge at them, they seem to disappear in a puff of blue smoke."
Xylotrupes socrates (Siamese rhinoceros beetle, or "fighting beetle"), male, on a banana leaf. This scarab beetle is particularly known for its role in insect fighting in Northern Laos and Thailand.
The orb-weaver spiders (family Araneidae) are the familiar builders of spiral wheel-shaped webs often found in gardens, fields and forests. The family is a large one, including over 2800 species in over 160 genera worldwide, making it the third largest known (behind Salticidae and Linyphiidae). The web has always been thought of as an engineering marvel.
The rose chafer (Cetonia aurata) is a reasonably large beetle (20 mm/¾ in long) that has metallic green coloration with a distinct V shaped scutellum, the small triangular area between the wing cases just below the thorax. Rose chafers are found over southern and central Europe and the southern part of the UK.
A Eusthenia species of stonefly. The order contains almost 3,500 known species, including the only known insects that are exclusively aquatic from birth to death. Stoneflies are believed to be one of the most primitive groups of Neoptera and are found worldwide, with the exception of Antarctica.
The cryptic mantis (Sibylla pretiosa) is a mantid native to southern Africa. Its common name comes from its ability to grow asymmetrically to match the vegetation of its environment. They have unusual leaf-like projections on the joint of their four walking legs and leaf-like wings, generally the only green portion of the insect's otherwise brown and mottled exoskeleton. Adult females grow to 5–6 cm (2.0–2.4 in) in length while the males are generally about 1 cm smaller.
Two flesh-flies (Sarcophaga ruficornis species pictured) mating. The life cycle of the saprophagic flesh-fly larvae has been well researched and is very predictable. Different species prefer bodies in different states of decomposition, which allows forensic entomologists to extrapolate the time of death.
The yellow fever mosquito (Aedes aegypti) is a species of mosquito known for its ability to spread yellow fever and dengue fever. The mosquito can be recognized by the white markings on its legs and a marking in the form of a lyre on its thorax. Though originally from Africa, the yellow fever mosquito can now be found in tropical regions around the world.
A European paper wasp (Polistes dominula) heating a bubble of regurgitated fluid in the sunlight. This is a common practice among many winged insects, and it is believed to be a way to facilitate digestion or to cool themselves off.
Papilio polymnestor, the blue Mormon, is a species of swallowtail butterfly found in southern India and Sri Lanka. It is a woodland species, often seen on forest paths and near streams. The larvae feed on trees in the family Rutaceae, such as citrus. Young larvae are green with white markings and position themselves on the upper surface of leaves, relying on their cryptic colouring, which resembles bird droppings, for protection. Older larvae seek less conspicuous locations, and have a unique habit of securing their balance by weaving silk on the substratum. This adult male P. polymnestor butterfly was photographed in the Indian state of Kerala.
The oriental hornet (Vespa orientalis) is a social insect in the family Vespidae, found in southern Europe, northeastern Africa, and western Asia. This worker, photographed in Sha'ar Poleg Reserve in Israel, is gathering nectar from a sea squill; adults will also feed on honeydew and fruit. They also capture insects such as grasshoppers, flies and honey bees to provide a diet high in protein for the colony's brood.
The European paper wasp (Polistes dominula or Polistes dominulus) is one of the most common species of wasps in the genus Polistes. Originally described in 1791 by German naturalist Johann Ludwig Christ as Vespa dominula, it is native to southern Europe and North Africa, as well as temperate parts of Asia as far east as China. It has also been introduced to Australia, New Zealand, South Africa and the Americas. The species is common and cosmopolitan due to its exceptional survival features, such as a productive colony cycle, a short development time and a higher ability to endure predator attacks. It generally lives in temperate, terrestrial habitats, such as chaparral, forest and grassland biomes.
This picture shows a young European paper wasp queen guarding her nest and eggs.
Two Melangyna viridiceps (called Common Hoverflies in Australia) mating in mid-air. The male, which can be identified by the eyes meeting at the top of its head, is on top. The term "hoverfly" refers to about 6,000 species of flying insects in the family Syrphidae. They are often seen hovering at flowers and are important pollinators.
The Meadow Argus (Junonia villida) is a species of butterfly native to Australasia. Its brown wings are each covered with two distinctive black and blue eyespots as well as white and orange marks that appear on the edge of the wings. Males and females are similar in appearance and size, with females being slightly larger.
A mole cricket, an insect belonging to the Gryllotalpidaefamily. Mole crickets are common insects, found on every continent except Antarctica, but because they are nocturnal and spend nearly all their lives underground in extensive tunnel systems, they are rarely seen. This specimen is likely to be Gryllotalpa brachyptera and is about 3.5 cm (1.4 in.) in size.
The light blue soldier crab (Mictyris longicarpus) inhabits beaches in the Indo-Pacific region. Soldier crabs filter sand or mud for microorganisms. They congregate during the low tide, and bury themselves in a corkscrew pattern during high tide, or whenever they are threatened.
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